Coleoptera
Scarabaeidae by Blaine Mathison
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A WORLDWIDE CHECKLIST OF THE TRIBES AND GENERA OF AEGIALIINAE, APHODIINAE, TERMITOTROGINAE, AULONOCNEMINAE, SCARABAEINAE, AND COPRINAE (COLEOPTERA: SCARABAEOIDEA: SCARABAEIDAE)

Blaine A. Mathison 1
Martin Hardy 2
Aleš Bezdek 3
Paul Schoolmeesters 4
January - 2001

Scarabaeoidea Latreille, 1902
=Lamellicornia Latreille, 1817

Scarabaeidae Latreille, 1802

Aegialiinae Laporte de Castelnau, 1840

Aegialiini Laporte de Castelnau, 1840
 =Silluviini Landin, 1949
 
Aegialia Latreille, 1807 44 spp; Europe to southeast Asia, North America
=Psammoporus Thomson, 1863
=Dimalia Mulsant et Rey, 1869
=Caelius Lewis, 1895
=Leptaegialia Brown, 1931
=Rhysothorax Bedel, 1911
=Anomalaegialia Borwon, 1931
=Anomalaegialia Brown, 1931
=Silluvia Landin, 1949
Micraegialia Brown, 1931 1 sp; Canada (Manitoba to British Columbia) and USA (Michigan to Washington)
Saprus Blackburn, 1904 4 spp; Australia and Tasmania
=Sapriniana Strand, 1917

Eremazini Stebnicka, 1977
Eremazus Mulsant, 1851 5 spp; Europe, Asia to northern India
=Millingenia Sharp, 1874
=Tolisus Sharp, 1874

Aphodiinae Leach, 1815

Aphodiini Leach, 1815
Acanthaphodius A. Schmidt, 1909 1 sp; Argentina, Chile
Anomalurobius Paulian, 1952 1 sp; Ivory Coast
Aphodius Illiger, 1798 1500 spp; cosmopolitan
=Aphodius Fabricius, 1833
=Cytoderhinus Seabra, 1909
=Digonorhinus Seabra, 1909
=Hemicyclorhinus Seabra, 1909
=Candezeollus Stebnicka et Howden, 1995
=Ammoecioides Bordat, 1999
Aphodoharmogaster Endrödi, 1983 1 sp; tropical Africa
Apsteiniella A. Schmidt, 1916 3 spp; Iraq, Central Asia
=Jacobsonia Koshantshikov, 1912
=Yacobsonia Sharp, 1913
Cephalocyclus Dellacasa, Gordon, et Dellacasa, 1998 7 spp; Guatemala, Mexico
Cnemargulus Semenov, 1903 3 spp; Saudi Arabia and Central Asia
Cnemisus Motschulsky, 1868 4 spp; southern Russia, Central Asia to Mongolia

=Ahermes Reitter, 1891
=Cnemargus Harold, 1866
=Cnemargus Motschulsky, 1845
Coptochirus Harold, 1859 10 spp; South Africa
Dialytellus Brown, 1929 2 spp; eastern North America
Dialytes Harold, 1869 4 spp; North America
Doaphius Balthasar, 1941 2 spp; tropical Africa
Euheptaulacus Dellacasa, 1983 10 spp; Europe, Asia Minor to Siberia
Exaphodius, Endrödi, 1960 1 sp; tropical Africa
Harmodactylus Péringuey, 1901 1 sp; South Africa
Heptaulacus Mulsant, 1842 7 spp; Europe and northern Africa
=Heptaulaculus Dellacasa et Baraud, 1978
Koreoxyomus Kim, 1996 1 sp; Korea
Lorditomaeus Péringuey, 1901 23 spp; tropical Africa
Macroretrus Péringuey, 1908 3 spp; Africa
Mothon Semenov et Medvedev, 1927 1 sp; Ukraine
Mozartius Nomura et Nakane, 1951 3 spp; Japan
Osmanius Branco et Baraud, 1988 3 spp; Iran, Libya, Turkey
Oxycorythus Solsky, 1876 1 sp; Central Asia
Oxyomus Stephens, 1839 27 spp; Europe to southeast Asia, North America (introduced)
=Amidorinus W. Koshantshikov, 1912
=Renaudius Balthasar, 1951
Paracoptochirus Balthasar, 1963 3 spp; Europe, Asia Minor
Psammodaphodius Endrödi, 1976 1 sp; South Africa
Sugrames Reitter, 1894 4 spp; Central Asia
Sybax Boheman, 1857 3 spp; tropical Africa
=Hypoplatys Harold, 1869
Turanella Semenov, 1905 1 sp; Central Asia
=Isochirus Reitter, 1892
Xenoheptaulacus Hinton, 1934 1 sp; Columbia, Panama, Venezuela
Xeropsamobeus Saylor, 1937 2 spp; USA (southern California)

Didactyliini Pittino, 1984
Didactylia d’Orbigny, 1896 28 spp; cosmopolitan
Glaraphodius Balthasar, 1941 1 sp; tropical Africa
Nataia Endrödi, 1961 1 sp; tropical Africa
Orocanthus Endrödi, 1973 1 sp; tropical Africa
Pseudomothon Pittino, 1984 1 sp; Saudi Arabia
Pseudoxyomus Petrovitz, 1962 1 sp; South Africa
Trigonoscelus Petrovitz, 1963 3 spp; Asia Minor to Pakistan

Proctophanini Stebnicka et Howden, 1995
Australaphodius Balthasar, 1942 2 spp; Africa, Australia, New Zealand, Tasmania
Drepanocanthus Péringuey, 1901 7 spp; South Africa
=Goiginus Endrödi, 1982
Harmogaster Harold, 1861 1 spp; South Africa, Australia (introduced)
=Odontaphodius A. Schmidt, 1907
Proctophanes Harold, 1861 8 spp; Australia and Tasmania
=Proctammodes Blackburn, 1892

Eupariini Lepeletier de Saint-Fargeau et Audinet-Serville, 1828
Afroataenius Petrovitz, 1969 1 sp; West Africa
Airapus Stebnicka et Howden, 1996 14 spp; southeast Asia and Pacific Islands, Australia
Anaetius Petrovitz, 1969 1 sp; Eritrea, Ethiopia
Annegialia Howden, 1971 1 sp; North America (Utah)
Aphotaenius Cartwright, 1952 4 spp; Brazil, Columbia, Mexico, eastern USA

Arupaia Stebnicka, 1999 1 sp; Brazil
Ataenioides Petrovitz, 1973 1 sp; Brazil
Ataeniopsis Petrovitz, 1973 12 spp; southern USA to South America, West Indies
Ataenius Harold, 1867 330 spp; cosmopolitan
=Auperia Chevrolat, 1864
=Auperia Duval, 1856
=Brancoataenius Paulian, 1979
=Euparia Burmeister, 1877
=Euparia Erichson, 1847
=Hexalus Mulsant et Rey, 1870
=Oxyomus Blanchard, 1843
=Pseudoammoecius A. Schmidt, 1913
=Stenataenius Chalumeau, 1982
Australammoecius Petrovitz, 1958 6 spp; Australia
Batesiana Chalumeau, 1983 1 sp; Panama
Bobiricola Endrödi, 1973 1 sp; Ghana
Bruchaphodius Martínez, 1952 3 spp; Argentina
Cartwrightia Islas, 1958 3 spp; tropical America
Cnematoplatys A. Schmidt, 1913 6 spp; Nepal to Thailand, south to Australia, New Guinea
Euparia Lepeletier de Saint-Fargeau et Audinet-Serville, 1828 3 spp; southern USA to South America, West Indies
Euparixia Brown, 1927 5 spp; southwestern USA to Central America and Cuba
Euparixoides Hinton, 1936 2 sp; Brazil, Panama
Euparotrix Stebnicka et Howden, 1996 1 sp; Lord Howe Island
Fletchmanniella Stebnicka, 1999 1 sp; Brazil
=Fletchmania Stebnicka, 1999
 Haroldiataenius Chalumeau, 1981 2 spp; Mexico
Iarupea Martínez, 1953 3 spp; Argentina, Brazil, Paraguay
=Iarupoides Chalumeau, 1984
Iguazua Stebnicka, 1997 1 sp; Argentina
Lomanoxoides Stebnicka, 1999 5 spp; Argentina, Bolivia, Brazil, Panama
=Lomanoxoides Chalumeau, 1984
Martineziella Chalumeau, 1986 7 spp; Argentina, Bolivia, Brazil, Chile, Cuba, Guyana, Mexico, Paraguay, southern USA, Korea (introduced?)
Myrmecaphodius Martínez, 1952 1 sp; tropical America
Napoa Stebnicka, 1999 1 sp; Ecuador
Nettelislasia Martínez, 1952 1 sp; Mexico
=Nettelia Islas, 1945
Notocaulus Quedenfeldt, 1884 13 spp; Africa
=Caulonotus Machatschke, 1955
=Fahraea Bergroth, 1888
Paraplesiataenius Chalumeau, 1992 3 spp; Central and South America
Parataenius Balthasar, 1961 5 spp; southeastern USA to South America, Portugal (introduced) and Australia (introduced)
=Brancoataenius Paulian, 1979
Passalaphodius Martínez, 1953 1 sp; Bolivia
Passaliolla Balthasar, 1945 7 spp; Central and South America
Phalangochaeta Martínez, 1952 25 spp; southern USA to South America, West Indies
Pseudataenius Brown, 1927 3 spp; eastern North America
Pseudosaprosites Balthasar, 1972 1 sp; Eritrea, Ethiopia
Saprositellus Balthasar, 1967 1 sp; Brazil
Saprosites Redtenbacher, 1857 100 spp; cosmopolitan
=Cylindropterus Motschulsky, 1863
Saprovisca Stebnicka, 1993 1 sp; Philippines
Selviria Stebnicka, 1999 1 sp; Brazil
Setylaides Stebnicka, 1994 4 spp; southeast Asia, Japan
Simogonius Harold, 1871 4 spp; Congo, Ethiopia

=Liparochirus Péringuey, 1901

Lomanoxiini Stebnicka, 1999
Lomanoxia Martínez, 1951 4 spp; tropical America

Psammodiini Mulsant, 1842
Aphodopsammobius Endrödi, 1964 5 spp; central Africa and Australia
Brindalus Landin, 1960 5 spp; Europe
Diastictus Mulsant, 1842 1 sp; Europe and western Asia
Geopsammodius Gordon et Pittino, 1992 3 spp; southeastern USA and French Guiana
Granulopsammodius Rakovi_, 1981 11 spp; Africa, Europe and Asia
Ingogius Endrödi, 1976 1 sp; South Africa
Leiopsammodius Rakovi_, 1981 38 spp; cosmopolitan
Neopsammodius Rakovi_, 1986 9 spp; North and Central America
Odontopsammodius Gordon et Pittino, 1992 14 spp; tropical America
Petrovitzius Rakovi_, 1979 2 spp; Japan, Madagascar
Psammodius Fallén, 1807 20 spp; Africa, Palearctic to Oriental region, North America (introduced)
=Psammobius Heer, 1841
Sicardia Reitter, 1896 2 spp; Algeria, Libya, Tunisia
=Bipammosus Normand, 1938
Tesarius Rakovi_, 1981 5 spp; Australia, western North America, England (introduced)
Trichiopsammobius Petrovitz, 1963 1 sp; Brazil, Venezuela

Rhyssemini Pittino et Mariani, 1986
Afrodiastictus Pittino et Mariani, 1986 5 spp; Fundu Isl., Ghana, Guinea, Ivory Coast, Mali, Senegal, Zanzibar, Zaire, South America
Bordatius Pittino et Mariani, 1986 4 spp; Morocco, South Africa
Messyrhus Rakovi_ et Král, 1997 1 sp; China
Myrhessus Balthasar, 1955 3 spp; Palearctic region
Mysarus Petrovitz, 1962 1 sp; South America
Neorhyssemus Gordon et Pittino, 1992 4 spp; South America
Neotrichiorhyssemus Rakovi_ et Král, 1997 14 spp; southeast Asia, Pacific Islands
Pararhyssemus Balthasar, 1955 1 sp; northern Africa to Kenya, Saudi Arabia to southwest Asia
Pittinius Rakovi_ et Král, 1997 1 sp; India, Nepal
Platytomus Mulsant, 1842 25 spp; North America, Africa, Europe to southeast Asia
Pleurophorus Mulsant, 1842 29 spp; Africa, Europe to southeast Asia, North America (introduced) south to Chile (introduced)
Psammorpha Stebnicka, 1994 1 sp; Christmas Island
Rhyssemodes Reitter, 1892 10 spp; northern Africa to Central Asia
Rhyssemorphus Clouët, 1900 5 spp; Madagascar
Rhyssemus Mulsant, 1842 200 spp; cosmopolitan
=Boucardius Petrovitz, 1967
Trichiorhyssemus Clouët, 1901 20 spp; southern USA to Central America, Africa to southeast Asia

Phycochini Rakovi_ et Král, 1997
Phycochus Broun, 1886 1 sp; New Zealand, Tasmania

Thinorycterini Semenov-Tyan-Shansky et Reichardt, 1925
Thinorhycter Semenov et Reichardt, 1925 6 spp; Middle Asia

Rhyparini A. Schmidt, 1910
Hadrorhyparus Howden, 1995 1 sp; Fiji
Rhyparus Westwood, 1843 50 spp; cosmopolitan (except North America)
=Antrisis Pascoe, 1866
=Ryparus Dejean, 1833

=Ryparus Westwood, 1843
Sybacodes Fairmaire, 1896 3 spp; Asia

Stereomerini Howden et Storey, 1992
Adebrattia Bordat et Howden, 1995 1 sp; Borneo, Malaysia
Australoxenella Howden et Storey, 1992 11 spp; Australia
Bruneixenus Howden et Storey, 1992 1 sp; Borneo, Brunei, Malaysia
Danielssonia Bordat et Howden, 1995 1 sp; Borneo, Malaysia
Pseudostereomera Bordat et Howden, 1995 1 sp; Borneo, Malaysia
Stereomera Arrow, 1905 1 sp; Singapore
Afroataenius Petrovitz, 1969 1 sp; West Africa
Airapus Stebnicka et Howden, 1996 14 spp; southeast Asia and Pacific Islands, Australia
Anaetius Petrovitz, 1969 1 sp; Eritrea, Ethiopia
Annegialia Howden, 1971 1 sp; North America (Utah)
Aphotaenius Cartwright, 1952 4 spp; Brazil, Columbia, Mexico, eastern USA

Arupaia Stebnicka, 1999 1 sp; Brazil
Ataenioides Petrovitz, 1973 1 sp; Brazil
Ataeniopsis Petrovitz, 1973 12 spp; southern USA to South America, West Indies
Ataenius Harold, 1867 330 spp; cosmopolitan
=Auperia Chevrolat, 1864
=Auperia Duval, 1856
=Brancoataenius Paulian, 1979
=Euparia Burmeister, 1877
=Euparia Erichson, 1847
=Hexalus Mulsant et Rey, 1870
=Oxyomus Blanchard, 1843
=Pseudoammoecius A. Schmidt, 1913
=Stenataenius Chalumeau, 1982
Australammoecius Petrovitz, 1958 6 spp; Australia
Batesiana Chalumeau, 1983 1 sp; Panama
Bobiricola Endrödi, 1973 1 sp; Ghana
Bruchaphodius Martínez, 1952 3 spp; Argentina
Cartwrightia Islas, 1958 3 spp; tropical America
Cnematoplatys A. Schmidt, 1913 6 spp; Nepal to Thailand, south to Australia, New Guinea
Euparia Lepeletier de Saint-Fargeau et Audinet-Serville, 1828 3 spp; southern USA to South America, West Indies
Euparixia Brown, 1927 5 spp; southwestern USA to Central America and Cuba
Euparixoides Hinton, 1936 2 sp; Brazil, Panama
Euparotrix Stebnicka et Howden, 1996 1 sp; Lord Howe Island
Fletchmanniella Stebnicka, 1999 1 sp; Brazil
=Fletchmania Stebnicka, 1999
Haroldiataenius Chalumeau, 1981 2 spp; Mexico
Iarupea Martínez, 1953 3 spp; Argentina, Brazil, Paraguay
=Iarupoides Chalumeau, 1984
Iguazua Stebnicka, 1997 1 sp; Argentina
Lomanoxoides Stebnicka, 1999 5 spp; Argentina, Bolivia, Brazil, Panama
=Lomanoxoides Chalumeau, 1984
Martineziella Chalumeau, 1986 7 spp; Argentina, Bolivia, Brazil, Chile, Cuba, Guyana, Mexico, Paraguay, southern USA, Korea (introduced?)
Myrmecaphodius Martínez, 1952 1 sp; tropical America
Napoa Stebnicka, 1999 1 sp; Ecuador
Nettelislasia Martínez, 1952 1 sp; Mexico
=Nettelia Islas, 1945
Notocaulus Quedenfeldt, 1884 13 spp; Africa
=Caulonotus Machatschke, 1955
=Fahraea Bergroth, 1888
Paraplesiataenius Chalumeau, 1992 3 spp; Central and South America
Parataenius Balthasar, 1961 5 spp; southeastern USA to South America, Portugal (introduced) and Australia (introduced)
=Brancoataenius Paulian, 1979
Passalaphodius Martínez, 1953 1 sp; Bolivia
Passaliolla Balthasar, 1945 7 spp; Central and South America
Phalangochaeta Martínez, 1952 25 spp; southern USA to South America, West Indies
Pseudataenius Brown, 1927 3 spp; eastern North America
Pseudosaprosites Balthasar, 1972 1 sp; Eritrea, Ethiopia
Saprositellus Balthasar, 1967 1 sp; Brazil
Saprosites Redtenbacher, 1857 100 spp; cosmopolitan
=Cylindropterus Motschulsky, 1863
Saprovisca Stebnicka, 1993 1 sp; Philippines
Selviria Stebnicka, 1999 1 sp; Brazil
Setylaides Stebnicka, 1994 4 spp; southeast Asia, Japan
Simogonius Harold, 1871 4 spp; Congo, Ethiopia

=Liparochirus Péringuey, 1901

Lomanoxiini Stebnicka, 1999
Lomanoxia Martínez, 1951 4 spp; tropical America

Psammodiini Mulsant, 1842
Aphodopsammobius Endrödi, 1964 5 spp; central Africa and Australia
Brindalus Landin, 1960 5 spp; Europe
Diastictus Mulsant, 1842 1 sp; Europe and western Asia
Geopsammodius Gordon et Pittino, 1992 3 spp; southeastern USA and French Guiana
Granulopsammodius Rakovi_, 1981 11 spp; Africa, Europe and Asia
Ingogius Endrödi, 1976 1 sp; South Africa
Leiopsammodius Rakovi_, 1981 38 spp; cosmopolitan
Neopsammodius Rakovi_, 1986 9 spp; North and Central America
Odontopsammodius Gordon et Pittino, 1992 14 spp; tropical America
Petrovitzius Rakovi_, 1979 2 spp; Japan, Madagascar
Psammodius Fallén, 1807 20 spp; Africa, Palearctic to Oriental region, North America (introduced)
=Psammobius Heer, 1841
Sicardia Reitter, 1896 2 spp; Algeria, Libya, Tunisia
=Bipammosus Normand, 1938
Tesarius Rakovi_, 1981 5 spp; Australia, western North America, England (introduced)
Trichiopsammobius Petrovitz, 1963 1 sp; Brazil, Venezuela

Rhyssemini Pittino et Mariani, 1986
Afrodiastictus Pittino et Mariani, 1986 5 spp; Fundu Isl., Ghana, Guinea, Ivory Coast, Mali, Senegal, Zanzibar, Zaire, South America
Bordatius Pittino et Mariani, 1986 4 spp; Morocco, South Africa
Messyrhus Rakovi_ et Král, 1997 1 sp; China
Myrhessus Balthasar, 1955 3 spp; Palearctic region
Mysarus Petrovitz, 1962 1 sp; South America
Neorhyssemus Gordon et Pittino, 1992 4 spp; South America
Neotrichiorhyssemus Rakovi_ et Král, 1997 14 spp; southeast Asia, Pacific Islands
Pararhyssemus Balthasar, 1955 1 sp; northern Africa to Kenya, Saudi Arabia to southwest Asia
Pittinius Rakovi_ et Král, 1997 1 sp; India, Nepal
Platytomus Mulsant, 1842 25 spp; North America, Africa, Europe to southeast Asia
Pleurophorus Mulsant, 1842 29 spp; Africa, Europe to southeast Asia, North America (introduced) south to Chile (introduced)
Psammorpha Stebnicka, 1994 1 sp; Christmas Island
Rhyssemodes Reitter, 1892 10 spp; northern Africa to Central Asia
Rhyssemorphus Clouët, 1900 5 spp; Madagascar
Rhyssemus Mulsant, 1842 200 spp; cosmopolitan
=Boucardius Petrovitz, 1967
Trichiorhyssemus Clouët, 1901 20 spp; southern USA to Central America, Africa to southeast Asia

Phycochini Rakovi_ et Král, 1997
Phycochus Broun, 1886 1 sp; New Zealand, Tasmania

Thinorycterini Semenov-Tyan-Shansky et Reichardt, 1925
Thinorhycter Semenov et Reichardt, 1925 6 spp; Middle Asia

Rhyparini A. Schmidt, 1910
Hadrorhyparus Howden, 1995 1 sp; Fiji
Rhyparus Westwood, 1843 50 spp; cosmopolitan (except North America)
=Antrisis Pascoe, 1866
=Ryparus Dejean, 1833

=Ryparus Westwood, 1843
Sybacodes Fairmaire, 1896 3 spp; Asia

Stereomerini Howden et Storey, 1992
Adebrattia Bordat et Howden, 1995 1 sp; Borneo, Malaysia
Australoxenella Howden et Storey, 1992 11 spp; Australia
Bruneixenus Howden et Storey, 1992 1 sp; Borneo, Brunei, Malaysia
Danielssonia Bordat et Howden, 1995 1 sp; Borneo, Malaysia
Pseudostereomera Bordat et Howden, 1995 1 sp; Borneo, Malaysia
Stereomera Arrow, 1905 1 sp; Singapore

Odochilini Rakovi_, 1987
Odochilus Harold, 1877 15 spp; southeast Asia, Pacific Islands

Odontolochini Stebnicka et Howden, 1996
Gongrolophus Stebnicka et Howden, 1996 1 sp; Australia
Loeblietus Endrödi, 1975 1 sp; Ivory Coast
Odontolochus A. Schmidt, 1916 18 spp; equatorial Africa and Australia
=Anodontolochus Paulian, 1942
=Odontoderus Clouët, 1900
Odontolytes Koshantshikov, 1916 3 spp; South America

Termitoderini Tangelder et Krikken, 1982
Termitaxis Krikken, 1970 1 sp; Peru
Termitoderus Mateu, 1966 2 spp; Angola, Chad, Namibia
Termitodiellus Nakane, 1961 1 sp; Micronesia
Termitodius Wasmann, 1894 10 spp; Borneo, Sulawesi, Sumatra, South America

Corythoderini A. Schmidt, 1910
Chaetopisthes Westwood, 1845 20 spp; India, Sri Lanka
Corythoderus Klug, 1845 3 spp; Chad, Ethiopia, India, Nigeria, Senegal, Sudan
=Eurycorythoderus Wasmann, 1918
=Stenocorythoderus Wasmann, 1918
Hemicorythoderus Tangelder et Krikken, 1982 1 sp; Zaire
Paracorythoderus Wasmann, 1918 3 spp; Namibia, South Africa
Termitopisthes Wasmann, 1918 7 spp; Burma, India

Termitotroginae Wasmann, 1918
Aphodiocopris Arrow, 1920 2 spp; Bangladesh, India
Termitotrox Reichensperger, 1915 5 spp; India, South Africa

Aulonocneminae Janssens, 1946
Ankaratrotrox Paulian, 1954 6 spp; Madagascar, Seychelles
Aulonocnemis Klug, 1838 40 spp; Comoros, Madagascar, South Africa
=Dialytoderus W. Koshantshikov, 1913
Manjarivolo Paulian, 1974 3 spp; Madagascar
Palnia Stebnicka, 1985 3 spp; China, India

Scarabaeinae Latreille, 1802

Canthonini Péringuey, 1900
Agamopus Bates, 1887 4 spp; Central and South America
Aleiantus Olsoufieff, 1947 20 spp; Madagascar
Amphistomus van Lansberge, 1871 21 spp; Australia, Melanesia

=Merodontus MacLeay, 1874
=Platyphymatia Waterhouse, 1874
Anachalcos Hope, 1837 8 spp; tropical Africa
=Chalconotus Reiche, 1841
Anisocanthon Martínez et Pereira, 1956 4 spp; tropical America
Anonthobium Paulian, 1984 6 spp; New Caledonia
Aphengoecus Péringuey, 1901 2 spp; South Africa
Apotolamprus Olsoufieff, 1947 8 spp; Madagascar
Aptenocanthon Matthews, 1974 3 spp; Australia
Arachnodes Westwood, 1837 69 spp; Madagascar
Aulacopris White, 1859 3 spp; Australia
Baloghonthobium Paulian, 1986 1 sp; New Caledonia
Bohepilissus Paulian, 1975 2 spp; South Africa
Boletoscapter Matthews, 1974 2 spp; Australia
Byrrhidium Harold, 1869 3 spp; Namibia, South Africa
=Elassocanthon Kolbe, 1908
Caeconthobium Paulian, 1984 1 sp; New Caledonia
Cambefortantus Paulian, 1986 1 sp; Madagascar
Canthochilum Chapin, 1934 13 spp; Cuba, Puerto Rico
Canthon Hoffmannsegg, 1817 146 spp; Western Hemisphere
=Coprobius Latreille, 1829
=Coeloscelis Reiche, 1841
=Paedohyboma Kolbe, 1893
=Glaphyrocanthon Martínez, 1948
=Peltecanthon Pereira, 1953
=Goniocanthon Pereira et Martínez, 1956
=Geocanthon Pereira et Martínez, 1956
=Nesocanthon Pereira et Martínez, 1956
=Boreocanthon Halffter, 1958
=Canthomoechus Periera et Martínez, 1959
=Trichocanthon Pereira et Martínez, 1959
Canthonella Chapin, 1930 14 spp; West Indies and South America
=Ipselissus Olsoufieff, 1935
Canthonidia Paulian, 1939 2 spp; South America
Canthonosoma MacLeay, 1871 3 spp; Australia
=Homodesmius Sharp, 1873
Canthotrypes Paulian, 1939 1 sp; Brazil
Cephalodesmius Westwood, 1841 3 spp; Australia
Circellium Latreille, 1825 1 sp; South Africa
Coproecus Reiche, 1841 1 sp; Australia
Cryptocanthon Balthasar, 1942 14 spp; Mexico to Brazil
Deltepilissus Periera, 1949 2 sp; Brazil
Deltochilum Eschscholtz, 1822 81 spp; southern USA to South America
=Anamnesis Vigors, 1826
=Hyboma Audinet-Serville, 1828
=Meghyboma Kolbe, 1893
Diorygopyx Matthews, 1974 8 spp; Australia
Endroedyolus Scholtz et Howden, 1987 1 sp; South Africa
Epactoides Olsoufieff, 1947 2 spp; Madagascar
Epilissus Reiche, 1841 1 sp; Madagascar
Epirinus Reiche, 1841 23 spp; Lesotho, South Africa
=Endrodyantus Cambefort, 1978
Eudinopus Burmeister, 1840 1 sp; Argentina, southern Brazil
=Stenodactylus Brullé, 1834
Falsignambia Paulian, 1987 1 sp; New Caledonia
Gyronotus van Lansberge, 1874 5 spp; South Africa, Tanzania
Hammondantus Cambefort, 1978 1 sp; Namibia

Hansreia Halffter et Martínez, 1977 1 sp; Central America
Haroldius Boucomont, 1914 20 spp; tropical Africa, southeast Asia
=Ponerotrogus Silveri, 1924
=Afroharoldius Janssens, 1949
=Phaedotrogus Paulian, 1985
Holocanthon Martinez et Pereira, 1956 1 sp; tropical America
Ignambia Heller, 1916 1 sp; Melanesia
Janssensantus Paulian, 1976 2 spp; Tanzania
Lambroma Sharp, 1873 3 spp; Australia
Lepanus Balthasar, 1966 26 spp; Australia, Melanesia, southeast Asia
Macropanelus Ochi, Kon et Araya, 1998 1 sp; Sumatra
Madaphacosoma Paulian, 1975 3 spp; Madagascar, Tanzania
Malagoniella Martínez, 1961 9 spp; Central and South America
Megathopa Eschscholtz, 1822 2 spp; Argentina, southern Brazil, Chile, Paraguay
=Glauconia Paulian, 1938
Megathoposoma Balthasar, 1939 1 sp; Mexico and Central America
Melanocanthon Halffter, 1958 4 spp; eastern North America
Mentophilus Laporte de Castelnau, 1840 2 spp; Australia
=Aulacium Reiche, 1841
Monoplistes van Lansberge, 1874 6 spp; Australia
Namakwanus Scholtz et Howden, 1987 1 spp; Namibia
Nanos Westwood, 1837 12 spp; Madagascar
Nesovinsonia Martínez et Pereira, 1959 1 sp; Mauritius
Odontoloma Boheman, 1857 20 spp; Africa
=Diaglyptus d’Orbigny, 1902
Oficanthon Paulian, 1985 1 sp; New Guinea
Onthobium Reiche, 1860 13 spp; New Caledonia
Outenikwanus Scholtz et Howden, 1987 1 sp; South Africa
Panelus Lewis, 1895 19 spp; Africa and Asia
Paracanthon Balthasar, 1938 4 spp; Brazil
Parachorius Harold, 1873 4 spp; southeast Asia
Paronthobium Paulian, 1984 1 sp; Melanesia
Peckolus Scholtz et Howden, 1987 2 spp; South Africa
Penalus Paulian, 1985 1 sp; New Guinea
Peyrierasantus Paulian, 1976 1 sp; Madagascar
Phacosoma Boucomont, 1914 10 spp; Melanesia, southeast Asia
Phacosomoides Martínez et Pereira, 1959 1 sp; Madagascar
Pseudarachnodes Lebis, 1953 3 spp; Madagascar
Pseudignambia Paulian et Pluot-Sigwalt, 1984 2 spp; Australia
Pseudocanthon Bates, 1887 8 spp; southern USA to South America
=Opiocanthon Paulian, 1947
Pseudonthobium Paulian, 1984 2 spp; New Caledonia
Pseudophacosoma Paulian, 1975 1 sp;
Pycnopanelus Arrow, 1931 3 spp; Namibia, Sudan, southeast Asia
Saphobiamorpha Brookes, 1944 1 sp; New Zealand
Saphobius Sharp, 1873 13 spp; New Zealand
Sauvagesinella Paulian, 1934 3 spp; Australia
Scybalocanthon Martínez, 1948 16 spp; Central and South America
Scybalophagus Martínez, 1953 6 spp; tropical America and South Africa
=Pseudoepirinus Ferreira, 1964
Sikorantus Paulian, 1976 1 sp; Madagascar
Sinapisoma Boucomont, 1928 1 sp; French Guiana
Sphaerocanthon Olsoufieff, 1947 23 spp; tropical Africa
Streblopus van Lansberge, 1874 2 spp; Brazil
Sylvicanthon Halffter et Martínez, 1977 5 spp; tropical America
Tanzanolus Scholtz et Howden, 1987 1 sp; Tanzania
Temnoplectron Westwood, 1841 15 spp; Australia, Melanesia

Tesserodon Hope, 1837 13 spp; Australia, Melanesia
Tetraechma Blanchard, 1843 2 spp; tropical America
Vulcanocanthon Pereira et Martinez, 1960 1 sp; South America
Xenocanthon Martinez, 1952 1 sp; Argentina, Brazil
Zonocopris Arrow, 1932 1 sp; Argentina, Brazil, Paraguay

Eucraniini Brullé, 1834
Anomiopsoides Blackwelder, 1944 8 spp; Argentina
=Anomiopsis Burmeister, 1861
Ennearabdus van Lansberge, 1874 1 sp; Argentina
Eucranium Brullé, 1834 7 spp; Argentina, Bolivia, Ecuador
=Anomiopsis Westwood, 1837
=Cyclodema Laporte de Castelnau, 1840
=Psammotrupes Guérin-Méneville, 1838
Glyphoderus Westwood, 1837 3 spp; Argentina

Eurysternini Vulcano, Martínez et Pereira, 1960
Eurysternus Dalman, 1824 26 spp; Mexico to South America
=Aeschrotes Audinet-Serville, 1828

Gymnopleurini Lacordaire, 1856
Allogymnopleurus Janssens, 1940 19 spp; Africa and Palearctic
Garreta Janssens, 1940 20 spp; Africa, southeast Asia
Gymnopleurus Illiger, 1803 52 spp; Africa, Europe to southeast Asia
Paragymnopleurus Shipp, 1897 13 spp; southeast Asia

Sisyphini Mulsant, 1842
Neosisyphus G. Müller, 1942 24 spp; Africa, Australia (introduced)
Nesosisyphus Vinson, 1946 4 spp; Mauritius
Sisyphus Latreille, 1807 30 spp; Africa, Europe to southeast Asia, Central America

Scarabaeini Latrielle, 1802
Drepanopodus Janssens, 1940 2 spp; Namibia, South Africa
Kheper Janssens, 1940 22 spp; Africa, Europe to southeast Asia
Pachylomera Kirby, 1828 2 sp; southern Africa
=Pachylomerus Bertoloni, 1849
Scarabaeus Linnaeus, 1758 112 spp; Africa and Madagascar, Mediterranean to Asia
=Actinophorus Creutzer, 1799
=Mnematium MacLeay, 1821
=Pachysoma MacLeay, 1821
=Sebasteos Westwood, 1847
=Mnematidium Ritsema, 1883
=Neateuchus Gillet, 1911
=Neomnematium Janssens, 1938
=Madateuchus Paulian, 1953
=Neopachysoma Ferreira, 1953
Sceliages Westwood, 1837 6 spp; southern Africa


Coprinae Kolbe, 1805

Coprini Kolbe, 1805
Catharsius Hope, 1837 90 spp; Africa and Asia
=Catharsiocopris Balthasar, 1967
Chalcocopris Burmeister, 1846 1 sp; Brazil
Copridaspidus Boucomont, 1920 1 sp; Africa

Copris Müller, 1764 160 spp; cosmopolitan (introduced into Australia and Hawaii)
=Palaeocopris Pierce, 1946
=Microcopris Balthasar, 1958
Coptodactyla Burmeister, 1846 13 spp; Australia, Melanesia
=Boucomontia Paulian, 1933
Dichotomius Hope, 1838 148 spp; southern USA to South America
=Homocopris Burmeister, 1842
=Selenocopris Burmeister, 1842
=Pinotus Erichson, 1847
=Brachycopris Haldeman, 1848
=Pseudoheliocopris Ferriera, 1970
Heliocopris Hope, 1837 49 spp; tropical Africa, southeast Asia
Holocephalus Hope, 1838 3 spp; southern Brazil, Paraguay
=Atrichius Gillet, 1907
Isocopris Pereira et Martínez 3 spp; Brazil
Litocopris Waterhouse, 1891 4 spp; Africa
Macroderes Westwood, 1876 13 spp; South Africa
Metacatharsius Paulian, 1939 61 spp; Africa
Ontherus Erichson, 1847 59 spp; Central and South America
Pseudocopris Ferreira, 1960 2 spp; Angola, Congo, Mozambique, Tanzania, Zambia, Zimbabwe
Pseudopedaria Felsche, 1904 2 spp; tropical Africa
Synapsis Bates, 1868 15 spp; Middle and southeast Asia
=Homalocopris Solsky, 1871
Thyregis Blackburn, 1904 4 spp; Australia
=Arrowianella Paulian, 1933
Xinidium Harold, 1869 3 spp; South Africa
=Parapinotus Harold, 1878

Ateuchini Montreuil, 1998
Anomiopus Westwood, 1842 29 spp; tropical America
Aphengium Harold, 1868 2 spp; southern South America
Ateuchus Weber, 1801 80 spp; southern and eastern USA to South America
=Choeridium Audinet-Serville, 1828
Bdelyropsis Pereira, Vulcano et Martínez, 1960 3 spp; Mexico to Venezuela
Bdelyrus Harold, 1869 23 spp; Central and South America
Canthidium Erichson, 1847 153 spp; tropical America
=Neocanthidium Martínez, Halffter et Pereira, 1964
Coptorhina Hope, 1830 18 spp; south-central Africa
=Frankenbergerius Balthasar, 1938
=Pseudocoptorhina Ferreira, 1954
Delopleurus Erichson, 1847 6 spp; Africa and India
Deltorhinum Harold, 1867 1 sp; Brazil
Demarziella Balthasar, 1961 14 spp; Australia
=Notopedaria Matthews, 1976
Hypocanthidium Balthasar, 1938 1 sp; Brazil
Onychothecus Boucomont, 1912 4 spp; southeast Asia
Paraphytus Harold, 1877 13 spp; Africa and Asia
=Xynophron Harold, 1880
=Maraxes Lewis, 1895
=Freyus Balthasar, 1958
Pedaria Laporte de Castelnau, 1832 34 spp; tropical Africa
Pedaridium Harold, 1868 11 spp; South America
Pseuduroxys Balthasar, 1938 1 sp; Ecuador
Pteronyx van Lansberge, 1874 1 sp; tropical America
Sarophorus Erichson, 1847 4 spp; South Africa
Scatimus Erichson, 1847 8 spp; Central and South America
Scatonomus Erichson, 1835 9 spp; Brazil

Trichillum Harold, 1868 19 spp; Central and South America
Uroxys Westwood, 1842 55 spp; tropical America

Onitini Hope, 1837
Acanthonitis Janssens, 1937 1 sp; Tanzania
Allonitis Janssens, 1936 1 sp; tropical Africa
Anonychonitis Janssens, 1950 1 sp; South Africa
Aptychonitis Janssens, 1937 3 spp; Kenya, Senegal
Bubas Mulsant, 1842 3 spp; Mediterranean, Australia (introduced)
Cheironitis van Lansberge, 1875 10 spp; Africa, Mediterranean to India
Gilletellus Janssens, 1937 2 spp; Mozambique, Zimbabwe
Heteronitis Gillet, 1911 5 spp; tropical Africa
Janssensellus Cambefort, 1976 1 sp; Cameroon
Kolbeellus Jacobson, 1906 1 sp; South Africa
Lophodonitis Janssens, 1938 1 sp; tropical Africa
Megalonitis Janssens, 1937 2 spp; Botswana, Mozambique
Neonitis Péringuey, 1901 2 spp; tropical Africa
Onitis Fabricius, 1798 153 spp; Africa, Mediterranean to southeast Asia, Australia (introduced) and USA (California, introduced)
Platyonitis Janssens, 1942 3 spp; tropical Africa
=Epionitis Balthasar, 1942
Pleuronitis van Lansberge, 1883 3 spp; western Africa
Pseudochironitis Ferreira, 1977 2 spp; tropical Africa
Tropidonitis Janssens, 1937 1 sp; Botswana, Zimbabwe

Phanaeini MacLeay, 1819
Bolbites Harold, 1868 1 sp; Argentina, southern Brazil, Paraguay, Uruguay
Coprophanaeus Olsoufieff, 1924 28 spp; Mexico to Argentina
=Megaphanaeus Olsoufieff, 1924
Dendropaemon Perty, 1830 25 spp; South America
Diabroctis Gistel, 1857 3 spp; South America
=Taurocopris Olsoufieff, 1924
Gromphas Brullé, 1834 4 spp; South America
Homalotarsus Janssens, 1932 1 sp; Brazil
Megatharsis Waterhouse, 1891 1 sp; Ecuador
Oruscatus Bates, 1870 2 spp; South America
Oxysternon Laporte de Castelnau, 1840 15 spp; southern Central America to South America
Phanaeus MacLeay, 1819 53 spp; southern and eastern USA to South America
=Lonchophorus Germar, 1824
=Onthurgus Gistel, 1857
Sulcophanaeus Olsoufieff, 1924 14 spp; southern Mexico to South America
Tetramereia Klages, 1907 1 sp; Brazil, Venezuela
=Eurypodea Klages, 1906
=Boucomontius Olsoufieff, 1924

Oniticellini d’Orbigny, 1916
Anoplodrepanus Simonis, 1981 2 spp; Jamaica
Cyptochirus Lesne, 1900 4 spp; Africa
Drepanocerus Kirby, 1828 25 spp; tropical Africa, China to southeast Asia
Drepanoplatynus Boucomont, 1921 1 sp; tropical Africa
Euoniticellus Janssens, 1953 17 spp; Africa, Europe to southeast Asia, Cuba, North America (introduced) and Australia (introduced)
Helictopleurus d’Orbigny, 1915 53 spp; Madagascar
Heterosyphus Paulian, 1975 1 sp; Madagascar
Liatongus Reitter, 1893 38 spp; Africa to Asia, western USA, Mexico and Central America, Australia (introduced)

Oniticellus Lepeletier de Saint-Fargeau et Audinet-Serville, 1828 9 spp; Africa to Asia
=Pseudoniticellus Kraatz, 1895
Paroniticellus Balthasar, 1963 1 sp; Asia Minor, Middle Asia
Scaptocnemis Péringuey, 1901 1 sp; Africa
Sinodrepanus Simonis, 1985 6 spp; China to southeast Asia
Tiniocellus Péringuey, 1900 6 spp; tropical Africa, southeast Asia
Tragiscus Klug, 1855 1 sp; Africa
Yvescambefortius Ochi et Kon, 1996 1 sp; Borneo, Sumatra, Malay Peninsula

Onthophagini Lacordaire, 1856
=Alloscelini Janssens, 1946
Alloscelus Boucomont, 1923 4 spp; Cameroon, Central African Republic, Gabon, Ivory Coast, Zaire
Amietina Cambefort, 1981 4 spp; Cameroon, Ivory Coast, Liberia, Malawi, Uganda
Anoctus Sharp, 1875 5 spp; southeast Asia
Caccobius Thomson, 1859 88 spp; Africa, Europe to southeast Asia
=Histeridium Motschulsky, 1859
=Caccophilus Jekel, 1872
Cambefortius Branco, 1989 9 spp; tropical Africa
Cassolus Sharp, 1875 7 spp; India, China south to Indonesia (Java, Sumatra, Celebes)
Cleptocaccobius Cambefort, 1984 24 spp; tropical Africa, southeast Asia
Cyobius Sharp, 1875 1 spp; Greater Sunda Islands
Diastellopalpus van Lansberge, 1886 23 spp; tropical Africa
Digitonthophagus Balthasar, 1959 18 spp; Africa, southeast Asia, Australia (introduced) and southern USA to South America (introduced)
Disphysema Harold, 1873 1 sp; India
Dorbignyolus Branco, 1989 1 sp; Ethiopia, Kenya, Somalia, Tanzania
Euonthophagus Balthasar, 1959 33 spp; Africa, Europe to southeast Asia
Eusaproecius Branco, 1989 7 spp; Cameroon, Gabon, Gambia, Kenya, Malawi, South Africa, Tanzania, Zaire, Zimbabwe
Heteroclitopus Péringuey, 1901 8 spp; Central African Republic, Guinea-Conakry, Natal, Nigeria, Senegal, Tanzania, Uganda, Zaire, Zimbabwe
Hyalonthophagus Palestrini et Giacone, 1988 10 spp; tropical Africa
Krikkenius Branco, 1991 1 sp; Kenya
Megaponerophilus Janssens, 1949 1 sp; Congo
Milichus Péringuey, 1901 8 spp; tropical Africa
Mimonthophagus Balthasar, 1963 2 spp; tropical Africa
Neosaproecius Branco, 1990 2 spp; Cameroon, Gabon, Liberia, Malawi, Zaire
Onthophagus Latreille, 1802 1800 spp; cosmopolitan
=Chalcoderus Erichson, 1848
=Monapus Erichson, 1848
=Psilax Erichson, 1848
=Gonocyphus van Lansberge, 1885
=Tauronthophagus Shipp, 1895
=Macropocopris Arrow, 1920
Phalops Erichson, 1847 37 spp; tropical Africa, Saudi Arabia to India
=Ephillopus Reitter, 1892
Pinacopodius Branco, 1991 1 sp; Kenya
Pinacotarsus Harold, 1875 1 spp; tropical Africa
Proagoderus van Lansberge, 1883 105 spp; Africa to India
Pseudosaproecius Balthasar, 1941 12 spp; tropical Africa
Stiptocnemis Branco, 1989 2 spp; Kenya, Tanzania
Stiptopodius Harold, 1871 9 spp; tropical Africa
=Saproecius Péringuey, 1901
Stiptotarsus Branco, 1989 2 spp; Cameroon, Gabon
Strandius Balthasar, 1935 17 spp; Africa to Asia
Sukelus Branco, 1992 1 sp; Kenya, Uganda
=Falcidius Branco, 1991

Tomogonus d’Orbigny, 1904 2 spp; tropical Africa
Walterantus Cambefort, 1977 1 sp; Zaire


1 1980 E. Harvard · Tempe, AZ 85283-2363 · USA · mathisb@hs.state.az.us
2 361 rue Ader · Beauport, Quebec · Canada G1C 5N6 · scarabs@sympatico.ca
3 ENTÚ AV_R · Branišovská 31 · 37005 Ceské Budejovice · Czech Republic · ales.bezdek@tix.bf.jcu.cz
4 Langeveldstraat, 23 · B-3020 Herent · Belgium · schoolp@online.be

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Last modified on Thursday, 25 July 2013